Esmée Winkel

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Selected publications
Book - Hyacintorchis

Hyacintorchis - Een nieuwe orchidee aan de Noordzee.
Author: Esmée Winkel. ISBN 978 90 5011 807 1, Order here

A contribution to Flora Batava, Uitgeverij Lannoo,
EAN: 9789401486668, Order here
Flora Batava cover
A contribution to Botanical Art Techniques. American Society of Botanical Artists. Authors: Carol Woodin, Robin Jess. ISBN-10 1604697903, ISBN-13 978-1604697902, Order here

Heukels' Flora van Nederland. Author: Leni Duistermaat, ISBN/EAN 9789001589561,

Wie wat bewaart. Author: Menno Schilthuizen. ISBN 9789000362493.

Van Onschatbare waarde, 200 jaar naturalis. Authors: Eulàlia Gassó Miracle and Tiny Monquil-Broersen. ISBN 9789462496279.

RHS Botanical Illustration - The Gold Medal Winners. Author: Charlotte Brooks. ISBN 978-1-78884-014-9, Order here

Flora Malesiana, Lamiaceae, Vol 23 - 2019, Series I - Seed Plants. Author: G.L.C. Bramley. ISBN 978-90-6519-019-2.

Out of the Woods: Celebrating Trees in Public Gardens. American Society of Botanical Artists & The New York Botanical Garden. Sku 978-0893279530.

Timber trees of Suriname, An identification Guide. Authors: Chequita R. Bhikhi, co-authors: Paul Maas, Jifke Koek-Noorman, Tinde van Andel. ISBN 9789460223914.

19th Annual International. American Society of Botanical Artists & The Horticultural Society of New York at New York Design Center. ISBN 978-0-9905616-5-1.

18th Annual International. American Society of Botanical Artists & The Horticultural Society of New York at New York Design Center. ISBN 978-0-9905616-3-7.

Description and drawings of the comparative morphology of mouth parts of various insects. 2013. Esmée Winkel. ISBN 978-90-818621-6-5.

Bloeiende Bollen, Blooming Bulbs. 2011. Vereniging botanisch kunstenaars Nederland. ISBN 978-90-818765-0-6.

Hunt Institute for Botanical Documentation. 2010. 13th International exhibition of botanical art & illlustration. ISBN 978-0-913196-84-7.

Siebolds bloemen tuin. 2010. Sieboldhuis. ISBN 2000000000718-8.

Contributions to scientific papers

Kasper P. Hendriks, Christiane Kiefer, Ihsan A. Al-Shehbaz, Félix Forest, Klaus Mummenhoff, Frederic Lens. 2023. Global Brassicaceae phylogeny based on filtering of 1,000-gene dataset. Current Biology. Read here.

J. Beyer, H.-J. Esser, M.C.M. Eurlings, P.C. van Welzen. 2023. Revision of the Genus Croton (Euphorbiaceae) in Sumatra (Indonesia). Blumea - Biodiversity Evolution and Biogeography of Plants 68(1). DOI:10.3767/blumea.2023.68.01.01. Read here.

Peter C. van Welzen, Esmée Winkel, Roderick W. Bouman. 2023. Parallel developments in floral adaptations to obligate moth pollination mutualism in tribe Phyllantheae (Phyllanthaceae). PhytoKeys 225: 165-198

De Wilde, W. J., & Duyfjes, B. (2022). The genus Canthium (Rubiaceae, Vanguerieae) in Thailand, with a note on the typification of the genus. Thai Forest Bulletin (Botany), 50(2), 161–180. Read here.

Jongkind, C.C.H. & O. Lachenaud. 2022. Novelties in African Apocynaceae. Candollea 77:17–51

Merijn A G de Bakker, Wessel van der Vos, Kaylah de Jager, Wing Yu Chung, Donald A Fowler, Esther Dondorp, Stephan N F Spiekman, Keng Yih Chew, Bing Xie, Rafael Jiménez, Constanze Bickelmann, Shigeru Kuratani, Radim Blazek, Peter Kondrashov, Marilyn B Renfree, Michael K Richardson, Selection on Phalanx Development in the Evolution of the Bird Wing, Molecular Biology and Evolution, 2021;, msab150. Read here.

C. A. J. Kreutz, J. C. Zuyderduyn. 2021. Hyacintorchis (Himantoglossum robertianum), een nieuwe orchideeënsoort voor Nederland. Gorteria – Dutch Botanical Archives  43, 2021: 070 – 088. Read here.

Fijridiyanto, I.A., Smets, E., Arifiani, D. 2020. Taxonomic revision of Dehaasia (Lauraceae) in Sumatra. Blumea 65, 2020: 167–175.

J. Hengstmengel. 2020. Emendations to tissue typology in discomycetes. Mycological Progress 19:543–558

R.-Y. Yu, P.C. van Welzen. 2020. A taxonomic revision of Trigonostemon (Euphorbiaceae) outside Malesia. Blumea 65,2020:25–52

R.-Y. Yu, E.M.G. Agoo, J.R. Callado, P.C. van Welzen. 2020. Taxonomic notes on Trigonostemon (Euphorbiaceae) in the Philippines. Blumea 65,2020:12–24.

Kusuma Wati R., Vugt R.R. van, Gravendeel B. 2018. A Linnaeus NG interactive key to the species of Glomera (Orchidaceae, Coelogyninae) from Southeast Asia. PhytoKeys (110) : 9-22.

Van Welzen, P.C. 2018. The genus Baliospermum (Euphorbiaceae) in Malesia. Blumea 63, 2018: 125–129.

Bouman, R.; van Welzen, P.C.; Sumail, S.; Echevarria, G.; Erskine, P.D.; van der EntBouman, A. 2018. Phyllanthus rufuschaneyi: a new nickel hyperaccumulator from Sabah (Borneo Island) with potential for tropical agromining. Botanical Studies (2018) 59:9.

Yu, R.-Y.; van Welzen, P.C. 2018. A taxonomic revision of Trigonostemon (Euphorbiaceae) in Malesia. Blumea 62, 2018: 179 – 229.

Heijkoop, M.; van Welzen, P.C. 2017. A revision of the genus Actephila (Phyllanthaceae) in the Malesian region. Blumea 62, 2017: 7– 25

Erkens, R.H.J., Oosterhof, J., Westra, L.Y.T., Maas, P.J.M. 2017. Revisions of Ruizodendron and Pseudephedranthus (Annonaceae) including a new species and an overview of most up-to-date revisions of Neotropical Annonaceae genera. PhytoKeys 86: 75–96.

Junikka, L.; Maas, P.J.M.; Maas-van de Kamer, H.; Westra, L.Y.Th. 2016. Revision of Oxandra (Annonaceae). Blumea 61, 2016: 215–266
Van der Ent, A.; van Balgooy, M.; van Welzen, P.C. 2016. Actephila alanbakeri (Phyllanthaceae): a new nickel hyperaccumulating plant species from localised ultramafic outcrops in Sabah (Malaysia). Van der Ent et al. Bot Stud  (2016) 57:6

Phyo Kay Khine, Stuart Lindsay, Christopher Fraser-Jenkins, Jürgen Kluge, Myint Kyaw, Peter Hovenkamp. 2016. Selliguea kachinensis (Polypodiaceae), a new fern species of uncertain affinity from Northern Myanmar. PhytoKeys 62: 73–81 (2016)

De Vogel, Winkel, E., van Vugt, R. 2016. Dendrobium cleistanthum and D. kruizingae, Two New Species of Section Crinifera from Papua New Guinea. Malesian Orchid J. 18: 108.

Junikka, L., Maas, P.J.M., Maas-van de Kamer, H., Westra, L.Y.T. 2016. Revision of Oxandra (Annonaceae). Blumea 61, 2016: 215-266.

Hoekstra PH, Wieringa JJ, Chatrou LW (2016) A nonet of novel species of Monanthotaxis (Annonaceae) from around Africa. PhytoKeys 69: 71-103.

Adema, F., Ohashi, H., Sunaro, B. 2016. Notes on Malesian Fabaceae (Leguminosae - Papilionoideae) 17. The genus Dalbergia. Blumea 61, 2016: 186-206.

Wiriadinata, H., Ohashi H., Adema, F. 2016. Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna. Blumea 61, 2016: 90-124.

Ottens-Treurniet, M.A.D. & Van Welzen, P.C. 2016. A revision of the Malesian genus BlumeodendronEuphorbiaceae). Blumea 61, 2016: 64–82.

Dirkse, G.M., Zonneveld, B.J.M., Duistermaat, L.H. 2015. Cardamine hamiltonii G.Don – Aziatische veldkers (Brassicaceae) in Nederland. Gorteria 37 (2014–2015): 64-70.

Maas, P.J.M., Westra, L.Y.T., Arias Guerrero, S. et al. 2015. Confronting a morphological nightmare: revision of the Neotropical genus Guatteria (Annonaceae). Blumea 60, 2015: 1-219.

Sulistyo B, Boos R, Cootes J, Gravendeel B. 2015. Dendrochilum hampelii (Coelogyninae, Epidendroideae, Orchidaceae) traded as ‘Big Pink’ is a new species, not a hybrid: evidence from nrITS, matK and ycf1 sequence data. PhytoKeys 56: 83-97. doi: 10.3897/phytokeys.56.5432

Van Welzen, P.C. & Winkel, E. 2015. A revision of Ostodes (Euphorbiaceae) in Malesia. Blumea 59, 2015: 185–190.

Karremans, A.P., Bogarín, D., Gravendeel, B. 2015. Specklinia lugduno-batavae (Pleurothallidinae: Orchidaceae), a new species in the S. digitalis group. Blumea 59, 2015: 180–184.

Van Welzen, P.C. & van Oostrum, A.F. 2015. Revision of the Malesian species of Dimorphocalyx  (Euphorbiaceae). Blumea 59, 2015: 191-201.

Veldkamp, J.F. 2015. Arundinella (Gramineae) in Malesia with notes on other taxa and on aluminium accumulation. Blumea 59, 2015: 167-179.

De Vogel, E., Winkel, E. and van Vugt, R. 2014. Dendrobium goodallianum, a new species of section Grastidium from Papua New Guinea. Malesian Orchid Journal 14: 87-90.

Sirichamorn, Y., Adema, F.A.C.B. & van Welzen, P.C. 2014. Derris solorioides (Fabaceae), a new limestone species with true-paniculate inflorescences from North-Central Thailand. Blumea 59, 2014: 98-102.

Hoekstra, P.H., Chatrou, L.W. & Jan J. Wieringa. 2014. A new species of Monanthotaxis from Gabon with a unique inflorescence type for Annonaceae. Phytotaxa 186(2): 106–11 2.

Dirkse, G.M., Reijerse, F.A.I., Duistermaat, L.H. 2014. De ontrafeling van in Nederland ingeburgerde Amerikaanse herfstasters (Symphyotrichum Nees subg. Symphyotrichum sect. Symphyotrichum, Asteraceae). Gorteria 37 (2014–2015): 1-24.

Stoops, E. & van Welzen, P.C. 2013. A revision of Ptychopyxis (Euphorbiaceae) in  southeast Asia. Nordic Journal of Botany 31: 94—112.

Tanawat Chaowasku & Paul J.A. Keßler. 2013. Phylogeny of Miliusa (Magnoliales: Annonaceae: Malmeoideae: Miliuseae), with descriptions of two new species from Malesia. European Journal of Taxonomy 54: 1-21, Fig. 3, 5.

Ed de Vogel, Rogier van Vugt, Malcolm Perry, Esmée Winkel & Eline Hoogendijk. 2013. Chelonistele maximae-reginae (Orchidaceae), an attractive new species of unknown provenance. Malesian Orchid Journal 12: 5-12. ISSN 1985-3912.

Kevin van den Hof, Thomas Marcussen, Ronald van den Berg, Eddy Weeda & Barbara Gravendeel. 2013. Phenotypic plasticity and nomenclature of Viola stagnina (V. persicifolia nom. ambig. props.). Gorteria 36: 61-84. ISSN 0017-2294.

Van Welzen, P.C. 2012. Five rare genera of Euphorbiaceae (sensu lato) in the Malay Archipelago: Alphandia, Ashtonia, Borneodendron, Cladogynos and Tapoides. Edinburgh Journal of Botany 69: 389—411. 

Pupulin, F., A.P. Karremans & B. Gravendeel. 2012. A reconsideration of the empusellous species of Specklinia (Orchidaceae: Pleurothallidinae) in Costa Rica. Phytotaxa 63: 1-20.

Van Welzen, P.C. & Forster, P.I. 2011. Picrodendraceae. In: Nooteboom, H. (ed.), Flora Malesiana ser. 1, 20: 45—61.

Van Welzen, P.C. 2011. Revision of Dicoelia (Phyllanthaceae; Euphorbiaceae s.l.). Blumea 56: 209—213.

Tanawat Chaowasku, Paul J.A. Keßler, Suwit Punnadee & Raymond W.J.M. van der Ham. 2011. Taxonomic novelties and pollen morphological study in the genus Neo-uvaria (Annonaceae). Phytotaxa 32: 27–42.

Arias Guerrero S., Welzen P.C. van 2011. Revision of Malesiana Endospermum (Euphorbiaceae) with notes on phylogeny and historical biogeography. Edinburgh Journal of Botany 68: 443-482.

Van Welzen, P.C. & Forster, P.I. 2010. A revision of Malesian Austrobuxus (Picrodendraceae/Euphorbiaceae s.l. subfam. Oldfieldioideae). Nordic Journal of Botany 28: 189—195

Hou, D., Savinov, I.A. & van Welzen, P.C. 2010. Celastraceae. In: Santisuk, T. & Larsen, K. (eds.), Flora of Thailand 10, 2: 141—198. The Forest Herbarium, Bangkok. 


Phd Theses

Dewi Pramanik. 2023. Evolution and development of flowers, fruits and inflorescences of Phalaenopsis and other orchid species. Naturalis Biodiversity Center and Leiden University.

Roderick Bouman. 2023. Disentangling a complex genus - Systematics, biogeography and bioactivity of the enus Phyllanthus L. and related genera of tribe Phyllantheae (Phyllanthaceae). Naturalis Biodiversity Center and Leiden University.

Abdulrokhman Kartonegoro. 2021. Systematics and Biogeography of the Dissochaeta alliance (Melastomataceae). Naturalis Biodiversity Center and Leiden University.

Rich Kusuma Wati. 2021. Systematics, epidermal defense and bioprospecting of wild orchids. Naturalis Biodiversity Center and Leiden University.

Ren-Yong Yu. 2019. A Monograph of the plant genus Trigonostemon Bluma. Naturalis Biodiversity Center and Leiden University.

Adam Karremans. 2015. Systematic, phylogenetic and pollination studies of Specklinia (Orchidaceae). Naturalis Biodiversity Center, Leiden University and Lankester Botanical Garden, Universidad de Costa Rica.

Tanawat Chaowasku. 2014. The Miliuseae revisited: Phylogenetic, taxonomic and palynological studies in a major clade of Annonaceae. Naturalis Biodiversity Center and Leiden University.

Abishkar Subedi. 2011. New species, pollinator interactions and pharmaceutical potential of Himalayan orchids. National Herbarium of the Netherlands, Leiden.

Hoang Van Sam. 2009. Uses and conservation of plant diversity in Ben En National park, Vietnam. National Herbarium of the Netherlands, Leiden. These drawings have also been published in: Hoang Van Sam, Baas, P. & Kessler, P.A.J. 2008. Plant Biodiversity in Ben En National Park, Vietnam. Agricultural Publishing House, Hanoi.

Kevin van den Hof. 2006. Scientific research regarding the evolutionary relationships of Viola stagnina and several closely related species. National Herbarium of the Netherlands, Leiden.